1. Fenner School of Environment and Society; The Australian National University; Canberra ACT 0200 Australia
2. Centre for Tropical Environmental and Sustainability Science (TESS), and School of Marine and Tropical Biology; James Cook University; Cairns Queensland 4878 Australia
3. School of Environmental and Forest Science; University of Washington; Seattle WA 98195 USA
4. Cary Institute of Ecosystem Studies; Millbrook NY 12545 USA
5. Department of Ecology and Evolutionary Biology; University of Connecticut, Storrs-Mansfield; CT 06269 USA
6. Department of Limnology; EBC-Uppsala University; 753 12 Uppsala Sweden