1. Key Laboratory of South China Agricultural Plant Molecular Analysis and Genetic Improvement South China Botanical Garden Chinese Academy of Sciences Guangzhou 510650 China
2. State Key Laboratory for Conservation and Utilization of Subtropical Agro‐Bioresources College of Horticulture South China Agricultural University Guangzhou 510642 China
3. China Key Laboratory of Biology and Germplasm Enhancement of Horticultural Crops in South China Ministry of Agriculture Guangzhou 510642 China
4. Key Laboratory of Plant Resources Conservation and Sustainable Utilization Guangdong Provincial Key Laboratory of Applied Botany South China Botanical Garden Chinese Academy of Sciences Guangzhou 510650 China