1. Department of Natural Resources and Environmental Management University of Hawaiʻi at Mānoa 1910 East‐West Road Honolulu HI 96822 U.S.A.
2. Department of Ecology and Evolutionary Biology Cornell University E145 Corson Hall Ithaca NY 14853 U.S.A.
3. USDA Forest Service, Pacific Southwest Research Station Institute of Pacific Islands Forestry 60 Nowelo Street Hilo HI 96720 U.S.A.
4. Department of Mathematics and Statistics University of Wyoming Laramie, 1000 E., University Avenue Laramie WY 82071 U.S.A.
5. Department of Environmental Sciences University of Puerto Rico PO Box 70377 San Juan PR 00936 U.S.A.