1. Laboratoire Evolution et Diversité Biologique – EDB, Université de Toulouse, UPS, 118 route de Narbonne, F‐31062, Toulouse, France
2. UMR 5174 – EDB, CNRS, Toulouse, France
3. Department of Environmental Science, Policy, and Management, University of California, 137 Mulford Hall, 94720 Berkeley, CA, USA