1. Department of Evolution, Ecology and Behaviour Institute of Integrative Biology Faculty of Health & Life Sciences University of Liverpool Liverpool L69 7ZB UK
2. Functional and Comparative Genomics Institute of Integrative Biology Faculty of Health & Life Sciences University of Liverpool Liverpool L69 7ZB UK
3. School of Biology University of St Andrews St Andrews KY16 9TH UK
4. Department of Biology University of York York YO10 5DD UK