1. Department of Biological and Environmental Science; University of Jyväskylä; Survontie 9 PO Box 35 Jyväskylä 40014 Finland
2. School of Biology; Dyers Brae; University of St Andrews; Greenside Place St Andrews Fife KY16 9TH UK
3. Animal and Plant Sciences; University of Sheffield; Western Bank Sheffield S10 2TN UK
4. School of Biosciences; University of Nottingham; Sutton Bonington Campus Loughborough LE12 5RD UK
5. Sven Lovén Centre for Marine Sciences-Tjärnö; University of Gothenburg; Strömstad SE 452 96 Sweden