1. Department of Ecology University of Granada Granada 18071 Spain
2. Area of Agriculture and Environment, Institute of Agricultural Research and Training Government of Andalusia Camino de Purchil s/n Granada 18004 Spain
3. Southern Swedish Forest Research Centre Swedish University of Agricultural Sciences Lund 23422 Sweden
4. Department of Agricultural and Environmental Science University of Milan 20133 Italy
5. Domingo Alcaraz‐Segura, Department of Botany University of Granada Granada 18071 Spain