1. Department of Fisheries; Islamic Azad University, Science and Research branch; Tehran PO Box 775/14515 Iran
2. Faculty of Marine Science; Tarbiat Modares University; PO Box 46414-356 Noor Iran
3. Department of Aquatic Animal Health, Faculty of Veterinary Medicine; University of Tehran; Tehran Iran
4. Department of Fisheries; Gorgan University of Agricultural Sciences and Natural Resources; Gorgan Iran