1. Institute of Plant Protection; Chinese Academy of Agricultural Science; No. 2 West Yuanmingyuan Road, Haidian District Beijing 100193 China
2. Institute of Plant and Environment Protection; Beijing Academy of Agriculture and Forestry Sciences; Beijing 100097 China
3. Huazhong Agricultural University; No. 1 Shizishan Street, Hongshan District Wuhan Hubei Province 430070 China
4. Department of Biological Chemistry and Crop Protection; Rothamsted Research; Harpenden Hertfordshire AL5 2JQ UK