1. Graduate School of Environmental Sciences; Gene Research Center; University of Tsukuba; 1-1-1 Tennodai Tsukuba 305-8572 Japan
2. RIKEN Center for Sustainable Resource Science; 1-7-22, Suehiro, Tsurumi Yokohama 230-0045 Japan
3. Boyce Thompson Institute for Plant Research; Ithaca NY 14853 USA
4. Partner State Key Laboratory of Agrobiotechnology; School of Life Sciences; The Chinese University of Hong Kong; Hong Kong China
5. U.S. Department of Agriculture/Agriculture Research Service; Robert W. Holley Centre for Agriculture and Health; Ithaca NY 14853 USA
6. Plant Biology Section; School of Integrative Plant Science; Cornell University; Ithaca NY 14853 USA