1. Department of Biology University of Victoria Victoria BC V8W 3N5 Canada
2. Aquatic Ecology and Evolution, Institute of Ecology and Evolution University of Bern Bern CH‐3012 Switzerland
3. Department of Fish Ecology and Evolution, Centre for Ecology, Evolution, and Biogeochemistry Swiss Federal Institute of Aquatic Science and Technology (EAWAG), Eawag ‐ Swiss Federal Institute of Aquatic Science and Technology Kastanienbaum CH‐6047 Switzerland
4. Natural History Museum Basel Basel CH‐4051 Switzerland
5. Howard Hughes Medical Institute, Stanford University School of Medicine Stanford California 94305 USA
6. Department of Developmental Biology Stanford University School of Medicine Stanford California 94305 USA
7. Friedrich Miescher Laboratory of the Max Planck Society Tübingen 72076 Germany
8. Earlham Institute Norwich NR4 7UZ United Kingdom
9. Department of Biological Sciences University of East Anglia Norwich NR4 7TJ United Kingdom