1. Wildlife Biology Program, Department of Ecosystem and Conservation Sciences, W.A. Franke College of Forestry and Conservation University of Montana Missoula Montana USA
2. Institute of Biodiversity, Animal Health and Comparative Medicine University of Glasgow Glasgow UK
3. NERC National Environmental Isotope Facility Scottish Universities Environmental Research Centre East Kilbride UK
4. SAERI – South Atlantic Environmental Research Institute Stanley Falkland Islands, UK