1. Quantitative Plant Ecology and Biodiversity Research Laboratory Department of Biology Faculty of Science Ferdowsi University of Mashhad Mashhad Iran
2. Instituto Multidisciplinar para el Estudio del Medio “Ramon Margalef” Universidad de Alicante Alicante Spain
3. Departamento de Ecología Universidad de Alicante Alicante Spain
4. Department of Range and Watershed Management Faculty of Natural Resources and Environment Ferdowsi University of Mashhad Mashhad Iran