1. INRA; UMR1202 BIOGECO; Cestas F-33610 France
2. Univ. Bordeaux; UMR1202 BIOGECO; Talence F-33400 France
3. Pernod Ricard Research Center; Créteil F-94000 France
4. Key Laboratory of Tropical Forest Ecology; Xishuangbanna Tropical Botanical Garden; Chinese Academy of Sciences; Mengla Yunnan 666303 China