1. Key Laboratory of Biodiversity and Biogeography Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China
2. Plant Germplasm and Genomics Center Germplasm Bank of Wild Species Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China
3. University of Chinese Academy of Sciences Beijing 100049 China
4. Royal Botanic Garden Edinburgh 20A Inverleith Row Edinburgh EH3 5LR UK
5. School of Biological Sciences Queen's University Belfast 97 Lisburn Road Belfast BT9 7BL UK