1. Department of Ecology and Environmental Science Umeå Plant Science Center, Umeå University Umeå SE‐90187 Sweden
2. CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Menglun Yunnan 666303 China
3. Forestry Research Institute of Sweden (Skogforsk) Sävar SE‐91833 Sweden
4. Center for Integrative Conservation & Yunnan Key Laboratory for Conservation of Tropical Rainforests and Asian Elephant, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences Menglun Yunnan 666303 China
5. Laboratory of Systematic Botany and Phytogeography, School of Biology Aristotle University of Thessaloniki Thessaloniki 54124 Greece
6. Graduate School of Agricultural Science Tohoku University Miyagi 989‐6711 Japan
7. National Engineering Research Center of Tree Breeding and Ecological Restoration, College of Biological Sciences and Technology Beijing Forestry University Beijing 100083 China