1. Institute of Paleontology Hebei GEO University 136 Huaiandonglu Shijiazhuang 050031 China
2. State Key Laboratory of Palaeobiology and Stratigraphy Nanjing Institute of Geology and Palaeontology, CAS Nanjing 210008 China
3. College of Life Sciences and Academy for Multidisciplinary Studies Capital Normal University 105 Xisanhuanbeilu, Haidian District Beijing 100048 China
4. Department of Paleobiology, National Museum of Natural History Smithsonian Institution Washington District of Columbia 20013‐7012 USA
5. Division of Entomology, Natural History Museum, and Department of Ecology & Evolutionary Biology University of Kansas Lawrence Kansas 66045 USA