Abstract
The description of a new fossil taxon Preheterobathmia grimaldiigen. nov. et sp. nov. is provided from Myanmar amber. The new taxon is tentatively assigned to the extant family Heterobathmiidae based on corresponding architecture of the female genitalia. It displays an interesting mixture of plesiomorphies and apomorphies, which are discussed in a comparison between Preheterobathmiagen. nov. and Heterobathmia. The fossil provides new information to the phylogeny and historical biogeography of ancestral Lepidoptera.