1. Innovative Center of Molecular Genetics and Evolution, School of Life Sciences Guangzhou University Guangzhou 510006 China
2. The Key Laboratory of Crop Genetics and Breeding of Hebei, Institute of Cereal and Oil Crops Hebei Academy of Agricultural and Forestry Sciences Shijiazhuang 050000 China
3. National Key Laboratory of Crop Genetics and Germplasm Enhancement, Jiangsu Collaborative Innovation Center for Modern Crop Production Nanjing Agricultural University Nanjing 210000 China
4. The Innovative Academy of Seed Design, Key Laboratory of Soybean Molecular Design Breeding, Northeast Institute of Geography and Agroecology the Chinese Academy of Sciences Harbin 150000 China