1. College of Marine Science and Technology Zhejiang Ocean University One Haidanan Road, Changzhi Island Zhoushan Zhejiang316022China
2. Department of Aquatic Science Faculty of Science Burapha University 169 Long‐Hard Bangsaen Road, Saensook Mueang Chonburi20131Thailand
3. State Key Laboratory of Genetic Resources and Evolution Kunming Institute of Zoology Chinese Academy of Sciences 21 Qingsong Road Kunming Yunnan650203China