1. State Key Laboratory of Microbial Resources; Institute of Microbiology; Chinese Academy of Sciences; Beijing 100101 China
2. University of the Chinese Academy of Sciences; Beijing 100049 China
3. School of Life Science; Liaoning University; Shenyang 110036 China
4. Center for Applied Geosciences; Eberhard Karls University Tuebingen; Tuebingen 72074 Germany
5. Department of Microbial Infection and Immunity; Department of Microbiology; Center for Microbial Interface Biology; The Ohio State University; Columbus OH 43210 USA