1. Department of Ecology and Evolutionary Biology; Yale University; 165 Prospect Street New Haven CT 06520 USA
2. Key Laboratory for Plant Biodiversity and Biogeography of East Asia; Kunming Institute of Botany; Chinese Academy of Sciences; Kunming 650201 Yunnan China
3. Department of Biostatistics; Yale School of Public Health; New Haven CT 06510 USA
4. Program in Microbiology; Yale University; New Haven CT 06520 USA