1. Division of Applied Life Sciences, Graduate School of Agriculture; Kyoto University; Kitashirakawa-Oiwake, Sakyo-ku Kyoto 606-8502 Japan
2. Faculty of Bioenvironmental Science; Department of Agriculture and Food Technology, Kyoto Gakuen University, 1-1 Nanjo-Ohtani; Sogabe Kameoka 621-8555 Japan
3. Department of Botany; Graduate School of Science, Kyoto University; Kitashirakawa-Oiwake, Sakyo-ku Kyoto 606-8502 Japan