1. State Key Laboratory of Agricultural Microbiology, College of Life Science and Technology Huazhong Agricultural University Wuhan 430070 China
2. Institute for Environmental Genomics, Department of Microbiology and Plant Biology University of Oklahoma Norman OK 73019 USA
3. Key Laboratory of Arable Land Conservation (Middle and Lower Reaches of Yangtze River), Ministry of Agriculture, College of Resources and Environment Huazhong Agricultural University Wuhan 430070 China