1. Department of Animal and Plant Sciences University of Sheffield Sheffield UK
2. Laboratoire Évolution and Diversité Biologique (EDB, UMR5174), CNRS, UPS, IRD Université de Toulouse Toulouse France
3. Claude E. Phillips Herbarium Delaware State University Dover Delaware
4. Genoscope CEA ‐ Institut de biologie François‐Jacob Evry Cedex France