1. Department of Biology and Biochemistry; University of Bath; Bath BA2 7AY UK
2. Milner Centre; University of Bath; Bath BA2 7AY UK
3. FAS Centre for Systems Biology; Harvard University; Cambridge MA 02138 USA
4. Center for Computational Biology and Bioinformatics; Department of Integrative Biology; The University of Texas; Austin TX 78712 USA
5. Department of Biology; Syracuse University; Syracuse NY 13244 USA