1. Department of Biosciences; Centre for Ecological and Evolutionary Synthesis; University of Oslo; PO Box 1066 Blindern N-0316 Oslo Norway
2. Centre for Geogenetics; Natural History Museum of Denmark; University of Copenhagen; Øster Voldgade 5-7 1350 Copenhagen K Denmark
3. Laboratoire AMIS; CNRS UMR 5288; Université de Toulouse, University Paul Sabatier (UPS); Toulouse France
4. McDonald Institute for Archaeological Research; University of Cambridge; Downing Street Cambridge CB2 3ER UK