1. Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology and Institute of Biodiversity School of Ecology and Environmental Science Yunnan University Kunming China
2. CAS Key Laboratory for Plant Diversity and Biogeography in East Asia Kunming Institute of Botany Chinese Academy of Sciences Kunming China
3. Department of Botany National Museum of Natural History Smithsonian Institution Washington DC USA
4. Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Mengla China