1. Co‐Innovation Center for Sustainable Forestry in Southern China Key Laboratory of Forest Genetics and Biotechnology of Ministry of Education Jiangsu Key Laboratory for Poplar Germplasm Enhancement and Variety Improvement Nanjing Forestry University Nanjing 210037 China
2. Department of Plant Biology Michigan State University East Lansing Michigan 48824 USA
3. Jiangsu Key Laboratory of Crop Genetics and Physiology Jiangsu Key Laboratory of Crop Genomics and Molecular Breeding Co‐Innovation Centre for Modern Production Technology of Grain Crops Key Laboratory of Plant Functional Genomics of the Ministry of Education Yangzhou University Yangzhou 225009 China
4. The State Key Laboratory of Crop Genetics and Germplasm Enhancement Jiangsu Collaborative Innovation Center for Modern Crop Production Nanjing Agricultural University Nanjing 210095 China
5. National Engineering Laboratory for Tree Breeding College of Bioscience and Biotechnology Beijing Forestry University Beijing 100083 China
6. Department of Horticulture Michigan State University East Lansing MI 48824 USA
7. Michigan State University AgBioResearch East Lansing MI 48824 USA