1. Department of Evolutionary Ecology; Museo Nacional de Ciencias Naturales; CSIC. José Gutiérrez Abascal 2 28006 Madrid Spain
2. Centre d’Etudes Biologiques de Chizé; UMR 7372 CNRS; Villiers-en-Bois France
3. Departamento de Ciencias Ambientales; Universidad de Castilla-La Mancha; Toledo Spain
4. Department of Tisza River Research; MTA Centre for Ecological Research; Danube Research Institute; Debrecen Hungary
5. Department of Conservation Biology; Estación Biológica de Doñana; Sevilla Spain