1. Key Laboratory of Biology and Genetic Improvement of Oil Crops, Ministry of Agriculture, Oil Crops Research Institute Chinese Academy of Agricultural Sciences Wuhan430062 HubeiChina
2. National Center for Gene Research, State Key Laboratory of Plant Molecular Genetics, CAS Center of Excellence in Molecular Plant Sciences, Institute of Plant Physiology and Ecology Chinese Academy of Sciences Shanghai 200233 China
3. Shanghai Key Laboratory of Plant Molecular Sciences, College of Life Sciences Shanghai Normal University Shanghai 200234 China