1. Centre d'Etudes Biologiques de Chizé; UMR 7372 CNRS-ULR 79360 Villiers en Bois France
2. Department of Biology; Virginia Tech; 4102 Derring Hall Blacksburg VA USA
3. Department of Evolutionary Zoology and Human Biology; University of Debrecen; Egyetem tér 1. Debrecen Hungary
4. Lendület Evolutionary Ecology Research Group; Plant Protection Institute; Centre for Agricultural Research; Hungarian Academy of Sciences; Herman Ottó út 15 H-1022 Budapest Hungary