1. 139 Hurst Street Oxford OX4 1HE UK
2. CIRAD; UR AGIRs; Ampandrianomby; BP 853; Antananarivo 101 Madagascar
3. Laboratoire des reptiles et amphibiens; Muséum national d'histoire naturelle; 25 rue Cuvier 75231 Paris cedex 05 France
4. Department of Archaeology & Natural History; Australian National University; Canberra ACT 0200 Australia
5. Life Sciences; University of Bristol; 24 Tyndall Avenue Bristol BS8 1TQ UK
6. Department of Biology; Earlham College; Richmond IN 47374 USA
7. Bird Group; Department of Life Sciences; Natural History Museum; Tring HP23 6AP UK
8. Wolfson College; Cambridge CB3 9BB UK