1. Key Laboratory of Plant Resources Conservation and Sustainable Utilization South China Botanical Garden Chinese Academy of Sciences Guangzhou China
2. University of Chinese Academy of Sciences Beijing China
3. South Subtropical Crops Research InstituteChinese Academy of Tropical Agriculture Sciences Zhanjiang China
4. Horticulture and Forestry Department Guangdong Ocean University Zhanjiang China
5. Innovation Academy of South China Sea Ecology and Environmental Engineering Chinese Academy of Sciences Guangzhou China