1. UMR BIOGER, INRA; AgroParisTech; Université Paris-Saclay; 78850 Thiverval-Grignon France
2. Christ Church; University of Oxford; St. Aldates Oxford OX1 1DP UK
3. Mathematical Institute; University of Oxford; Andrew Wiles Building, Radcliffe Observatory Quarter, Woodstock Road Oxford OX2 6GG UK
4. Department of Zoology; University of Oxford; South Parks Road Oxford OX1 3PS UK