1. Evolutionary Biology and Ecology CP160/12, Faculté des Sciences Université Libre de Bruxelles (ULB) Brussels Belgium
2. Herbarium de l'Université Libre de Bruxelles (BRLU) CP 265 Brussels Belgium
3. Okinawa Institute of Science and Technology (OIST) Onna‐son Okinawa Japan
4. Missouri Botanical Garden, Africa & Madagascar Department St. Louis Missouri USA
5. Meise Botanic Garden Meise Belgium