1. Department of Biology; Faculty of Science; Ferdowsi University of Mashhad; Mashhad 9177 9489 74 Iran
2. Research Department of Zoological Innovations; Institute of Applied Zoology; Faculty of Science; Ferdowsi University of Mashhad; Mashhad 9177 9489 74 Iran
3. Department of Social Sciences; Oxford Brookes University; Oxford OX3 0BP UK
4. Department of Ecology and Evolution; University of Lausanne; Biophore Building; CH-1015 Lausanne Switzerland