1. 1 Department of Biology, Faculty of Sciences Ferdowsi University of Mashhad Mashhad Iran
2. 6 E-mail: aliabadi@um.ac.ir
3. 2 Oxford Brookes University School of Social Sciences and Law, Department of Anthropology and Geography OX3 0BP Oxford United Kingdom
4. 3 Department of Computer Engineering, Faculty of Engineering Ferdowsi University of Mashhad Mashhad Iran
5. 4 Naturalis Biodiversity Center Darwinweg 2 2333 CR Leiden The Netherlands
6. 5 Zoological Institute Technical University Braunschweig Mendelssohnstr. 4 38106 Braunschweig Germany