1. Department of Animal Ecology and Tropical Biology Biocenter University of Würzburg Würzburg Germany
2. Bavarian Forest National Park Grafenau Germany
3. Chair for Strategic Landscape Planning and Management Technical University of Munich Freising Germany
4. Chair for Forest Growth and Yield Science School of Life Sciences Weihenstephan Technical University of Munich Freising Germany
5. Terrestrial Ecology Research Group Department of Ecology and Ecosystem Management Technical University of Munich Freising Germany