1. Krasnoyarsk Scientific Center, Siberian Branch of Russian Academy of Sciences (SB RAS), Krasnoyarsk 660036, Russia
2. V.N. Sukachev Institute of Forest, Siberian Branch of Russian Academy of Sciences (SB RAS), Krasnoyarsk 660036, Russia
3. Department of Ecology and Nature Management, Siberian Federal University, Krasnoyarsk 660041, Russia
4. Institute of Systematics and Ecology of Animals, Siberian Branch of Russian Academy of Sciences (SB RAS), Novosibirsk 630091, Russia