1. Research Centre for Ecological Change, Organismal and Evolutionary Biology Research Programme, Faculty of Biological and Environmental Sciences, Univ. of Helsinki Helsinki Finland
2. Dept of Ecology and Evolutionary Biology, Yale Univ. New Haven CT USA
3. Dept of Mathematics and Statistics, Faculty of Science, Univ. of Helsinki Helsinki Finland
4. Dept of Geosciences and Geography, Faculty of Science, Univ. of Helsinki Helsinki Finland