1. State Key Laboratory for Biology of Plant Diseases and Insect Pests, Institute of Plant ProtectionChinese Academy of Agricultural Sciences Beijing 100193 China
2. College of Plant ProtectionNanjing Agricultural University Nanjing 210095 China
3. Rothamsted Research, West Common, Harpenden Hertfordshire AL5 2JQ U.K.
4. National Animal Husbandry ServiceMinistry of Agriculture and Rural Affairs Beijing 100125 China
5. Centre for Ecology and Conservation, and Environment and Sustainability InstituteUniversity of Exeter Cornwall TR10 9FE U.K.