1. German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig Leipzig Germany
2. CEFE, Univ. de Montpellier, CNRS, EPHE, IRD, Univ. Paul Valéry Montpellier 3 Montpellier France
3. Centre for Mathematical Modelling of Infectious Diseases, London School of Hygiene and Tropical Medicine London UK