1. Groningen Inst. for Evolutionary Life Sciences, Univ. of Groningen; Box 11103, NL-9700 Groningen CC the Netherlands
2. CSIRO, Centre for Australian National Biodiversity Research; GPO Box 1600, Canberra ACT 2601 Canberra Australia
3. Theoretical Ecology Lab, Center for Advanced Studies of Blanes, CEAB-CSIC; Spain
4. Dept of Biology; Univ. of British Columbia- Okanagan; BC Canada