1. Department of Veterinary Clinical Sciences, Jockey Club College of Veterinary Medicine and Life Sciences City University of Hong Kong Hong Kong SAR China
2. Behavioral Ecology Group, Section for Ecology and Evolution, Department of Biology University of Copenhagen Copenhagen Denmark
3. Department of Infectious Diseases and Public Health Jockey Club College of Veterinary Medicine and Life Sciences, City University of Hong Kong Hong Kong SAR China
4. Centre for Animal Health and Welfare Jockey Club College of Veterinary Medicine and Life Sciences, City University of Hong Kong Hong Kong SAR China