1. Division of Biology and Conservation Ecology; Manchester Metropolitan University; Manchester M1 5GD United Kingdom
2. Kenya Wildlife Service; Samburu Research Centre; P.O. Box 53 Maralal Kenya
3. Blond McIndoe Laboratories; University of Manchester, Oxford Road; Manchester M13 9PT United Kingdom
4. International Livestock Research Institute; ILRI Kenya; P.O. Box 30709 Nairobi 00100 Kenya
5. Institute of Integrative Biology; Biosciences Building; Crown Street; University of Liverpool; Liverpool L69 7ZB United Kingdom
6. Kenya Wildlife Service; P.O. Box 40241-00100 Nairobi Kenya
7. Department of Biological and Environmental Science; FI-40014; University of Jyväskylä Finland