1. School of Biological Sciences; Queen's University Belfast; Belfast BT 9 7BL UK
2. Ecology and Evolutionary Biology; University of California Irvine; Irvine CA 92697 USA
3. School of Biology and Environmental Science; University College Dublin; Dublin Ireland
4. School of Natural Sciences; Department of Zoology; Trinity College Dublin; Dublin Ireland
5. Trinity Centre for Biodiversity Research; Trinity College Dublin; Dublin Ireland