1. Department of Wildland Resources and the Ecology Center Utah State University Logan UT 84322 USA
2. Centre for Excellence in Environmental Decisions School of Biological Sciences The University of Queensland Brisbane Qld 4072 Australia
3. Evolutionary Demography Laboratory Max Planck Institute for Demographic Research Konrad‐Zuße straße 1 Rostock DE‐18057 Germany
4. Department of Zoology School of Natural Sciences Trinity College Dublin Dublin 2 Ireland