1. Department of Pathology; University of Cambridge; Addenbrooke's Hospital; Cambridge UK
2. Division of Signal Transduction; Beth Israel Deaconess Medical Center and Department of Systems Biology; Harvard Medical School; Boston MA USA
3. Department of Histopathology; Institute of Orthopaedics and Musculoskeletal Science; Royal National Orthopaedic Hospital NHS Trust; University College London; Stanmore Middlesex UK
4. Division of Pathology; University of Edinburgh; Institute of Genetics & Molecular Medicine; Western General Hospital; Edinburgh UK