1. The Institute of Statistical Mathematics 10‐3 Midori‐cho Tachikawa Tokyo190‐8562 Japan
2. Department of Fish Ecology and Evolution Center for Ecology, Evolution & Biogeochemistry Swiss Federal Institute of Aquatic Science and Technology 6047 Kastanienbaum Switzerland
3. Graduate School of Agricultural and Life Sciences The University of Tokyo 1‐1‐1 Yayoi, Bunkyo‐ku Tokyo113‐8657 Japan